On re-fitting the wheel we did notice that you can tighten all nuts by hand, wiggle the wheel a bit, then tighten them some more and you can do this several times, so they are quite a finicky fit. This suggests that if you lowered all of the car's weight onto the wheels before fully tightening the wheel nuts there is a chance that you would not be able to torque them up properly with a typical cross style wheel brace, even though they appear to be very tight.
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